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{{Short description|Genus of flowering plants}}
{{Other uses}}
{{Other uses}}
{{Automatic taxobox
{{Taxobox
| image = Perilla.jpg
| image = Perilla.jpg
⚫
| image_caption = ''[[Perilla frutescens]]''
| image_alt =
| display_parents = 2
⚫
| image_caption = ''[[Perilla frutescens]]''
| taxon = Perilla (plant)
| regnum = [[Plantae]]
| unranked_divisio = [[Angiosperms]]
| authority = [[Carl Linnaeus|L.]]
| unranked_classis = [[Eudicots]]
| unranked_ordo = [[Asterids]]
| ordo = [[Lamiales]]
| familia = [[Lamiaceae]]
| genus = '''''Perilla'''''
| genus_authority = [[Carl Linnaeus|L.]]
}}
}}


'''''Perilla''''' is a genus consisting of one major Asiatic crop species ''[[Perilla frutescens]]'' and a few wild species in nature belonging to the mint family, [[Lamiaceae]]. The genus encompasses several distinct [[Variety (botany)|varieties]] of Asian [[herb]], [[seed]], and [[vegetable]] [[crop]], including ''[[Perilla frutescens|P. frutescens]]'' (deulkkae) and [[Perilla frutescens var. crispa|''P. frutescens'' var. ''crispa'']] (shiso).<ref name="Nitta">{{Cite journal|last=Nitta|first=Miyuki|last2=Lee|first2=Ju Kyong|last3=Ohnishi|first3=Ohmi|date=2003|title=AsianPerilla crops and their weedy forms: Their cultivation, utilization and genetic relationships|url=https://link.springer.com/article/10.1663/0013-0001(2003)057[0245:APCATW]2.0.CO;2|journal=Economic Botany|volume=57|issue=2|pages=245–253|doi=10.1663/0013-0001(2003)057[0245:APCATW]2.0.CO;2|issn=0013-0001}}</ref> The [[genus]] name ''Perilla'' is also a frequently employed common name ("perilla"), applicable to all varieties.<ref name="Vaughan">{{Cite book|url=https://books.google.com/books?id=2g99Q6-RSdAC&pg=PT340|title=The New Oxford Book of Food Plants|last=Vaughan|first=John|last2=Geissler|first2=Catherine|publisher=[[Oxford University Press]]|year=2009|isbn=9780199549467|edition=2nd|location=Oxford|pages=340}}</ref><ref name="Staples">{{Cite book|url=https://books.google.com/books?id=jbsdYHEJiDIC&pg=PA82|title=Ethnic Culinary Herbs: A Guide to Identification and Cultivation in Hawaiʻi|last=Staples|first=George W.|last2=Kristiansen|first2=Michael S.|publisher=[[University of Hawaiʻi Press]]|year=1999|isbn=9780824820947|location=Honolulu|pages=82}}</ref> Perilla varieties are [[Cross-fertilization|cross-fertile]] and intra-specific [[Hybrid (biology)|hybridization]] occurs [[Nature|naturally]].<ref name="Nitta" /> Some varieties are considered [[Invasive plant|invasive]].<ref name="NPS">{{Cite web|url=https://www.nps.gov/plants/ALIEn/pubs/midatlantic/pefr.htm|title=Beefsteak Plant (Perilla frustescens)|date=11 November 2010|website=[[National Park Service]]|publisher=[[U. S. Department of the Interior]]|access-date=4 December 2016}}</ref>
'''''Perilla''''' is a genus consisting of one major Asiatic crop species ''[[Perilla frutescens]]'' and a few wild species in nature belonging to the mint family, [[Lamiaceae]]. The genus encompasses several distinct [[Variety (botany)|varieties]] of Asian [[herb]], [[seed]], and [[vegetable]] [[crop]], including ''[[Perilla frutescens|P. frutescens]]'' (deulkkae) and [[Perilla frutescens var. crispa|''P. frutescens'' var. ''crispa'']] (shiso).<ref name="Nitta">{{Cite journal|last1=Nitta|first1=Miyuki|last2=Lee|first2=Ju Kyong|last3=Ohnishi|first3=Ohmi|date=2003|title=AsianPerilla crops and their weedy forms: Their cultivation, utilization and genetic relationships|journal=Economic Botany|volume=57|issue=2|pages=245–253|doi=10.1663/0013-0001(2003)057[0245:APCATW]2.0.CO;2|s2cid=30209741 |issn=0013-0001}}</ref> The [[genus]] name ''Perilla'' is also a frequently employed common name ("perilla"), applicable to all varieties.<ref name="Vaughan">{{Cite book|url=https://books.google.com/books?id=2g99Q6-RSdAC&pg=PT340|title=The New Oxford Book of Food Plants|last1=Vaughan|first1=John|last2=Geissler|first2=Catherine|publisher=[[Oxford University Press]]|year=2009|isbn=9780199549467|edition=2nd|location=Oxford|pages=340}}</ref><ref name="Staples">{{Cite book|url=https://books.google.com/books?id=jbsdYHEJiDIC&pg=PA82|title=Ethnic Culinary Herbs: A Guide to Identification and Cultivation in Hawaiʻi|last1=Staples|first1=George W.|last2=Kristiansen|first2=Michael S.|publisher=[[University of Hawaiʻi Press]]|year=1999|isbn=9780824820947|location=Honolulu|pages=82}}</ref> Perilla varieties are [[Cross-fertilization|cross-fertile]] and intra-specific [[Hybrid (biology)|hybridization]] occurs [[Nature|naturally]].<ref name="Nitta" /> Some varieties are considered [[Invasive plant|invasive]].<ref name="NPS">{{Cite web|url=https://www.nps.gov/plants/ALIEn/pubs/midatlantic/pefr.htm|title=Beefsteak Plant (Perilla frustescens)|date=11 November 2010|website=[[National Park Service]]|publisher=[[U. S. Department of the Interior]]|access-date=4 December 2016}}</ref>


== Taxa and synonyms ==
==Taxa and synonyms==
The classification of ''Perilla'' is confused, partly because botanists struggled with distinguishing the two distinct [[cultigen]]s as different species or variations.<ref name="Nitta" /><ref name="Yu">{{Cite book|url=https://books.google.com/books?id=M5A71gsWUskC&pg=PA1|title=Perilla: The Genus Perilla|publisher=[[CRC Press]]|year=1997|isbn=9789057021718|editor-last=Yu|editor-first=He-ci|location=London|pages=1–3|editor-last2=Kosuna|editor-first2=Kenichi|editor-last3=Haga|editor-first3=Megumi}}</ref> Until a few decades ago, [[Perilla frutescens var. crispa|''P. frutescens'' var. ''crispa'']] was regarded as a species in its own right, distinct from [[Perilla frutescens|''P. frutescens'']], although it was well established that these types readily cross-pollinate.<ref name="Heibonsha">{{Cite book|title=Sekai dai-hyakka jiten|title-link=Heibonsha World Encyclopedia|publisher=Heibonsha|year=1969|editor-last=Satake|editor-first=Yoshisuke|volume=10|location=Tōkyō|pages=246-207|script-title=ja:世界大百科事典|trans-title=Heibonsha World Encyclopedia|chapter=shiso|script-chapter=ja:しそ|orig-year=1968|editor-last2=Nishi|editor-first2=Sadao|editor-last3=Motoyama|editor-first3=Tekishū}}</ref> An early example of dividing the two cultigens into different species is found in [[Jinzō Matsumura|Matsumura]]'s 1884 nomenclature book, where the synonym ''P. arguta'' [[Benth.|<small>Benth.</small>]] is applied to [[Perilla frutescens var. crispa|''P. frutescens'' var. ''crispa'']], and the synonym ''P. ocymoides'' [[Carl Linnaeus|<small>L.</small>]] was applied to [[Perilla frutescens|''P.'' frutescens]].<ref name="Matsumura">{{Cite book|url=https://books.google.com/books?id=IOIlAQAAMAAJ&pg=PA136|title=Nippon shokubutsumeii|last=Matsumura|first=Jinzō|publisher=Z.P. Maruya|year=1884|location=Tōkyō|pages=136|script-title=ja:日本植物名彙|trans-title=Nomenclature of Japanese plants in Latin, Japanese and Chinese|author-link=Jinzō Matsumura}}</ref><ref name="Roecklein">{{Cite book|url=https://books.google.com/books?id=C8eRalwPua0C&pg=PA349|title=A Profile of Economic Plants|publisher=[[Transaction Publishers]]|year=1987|isbn=9780887381676|editor-last=Roecklein|editor-first=John C.|location=New Brunswick, U.S.A|pages=349|language=English|editor-last2=Leung|editor-first2=PingSun}}</ref><ref name="Kays">{{Cite book|url=https://books.google.com/books?id=9c7LggdxjBcC&pg=PA677|title=Cultivated Vegetables of the World:: A Multilingual Onomasticon|last=Kays|first=S. J.|publisher=[[Wageningen Academic Publishers]]|year=2011|isbn=9789086861644|location=Wageningen|pages=180–181, 677–678}}</ref><ref name="Blaschek">{{Cite book|url=https://books.google.com/books?id=XGXnVRiu3zoC&pg=PA328|title=Hagers Handbuch der Pharmazeutischen Praxis|publisher=[[Gabler Wissenschaftsverlage]]|year=1998|isbn=9783540616191|editor-last=Blaschek|editor-first=Wolfgang|edition=3|location=Berlin|pages=328-|language=de|editor-last2=Hänsel|editor-first2=Rudolf|editor-last3=Keller|editor-first3=Konstantin|editor-last4=Reichling|editor-first4=Jürgen|editor-last5=Rimpler|editor-first5=Horst|editor-last6=Schneider|editor-first6=Georg}}</ref> The species name ''P. ocymoides'' or ''P. ocimoides'' has historically been used to denote [[Perilla frutescens var. crispa|''P. frutescens'' var. ''crispa'']], especially by the Japanese,{{efn|e.g. occurs in Heibonsha 1964 Encyclopedia, though the genus name is misspelt}}<ref name="Heibonsha" /> therefore it should not be considered an interchangeable synonym for either cultigen. Recent genetic research confirms that the cultigens are of a common gene pool, corroborating the [[taxonomist]]s' claim for consolidating the two crops into one species.<ref name="Ito">{{Cite book|title=Phytochemistry Research Progress|last=Ito|first=Michiho|publisher=[[Nova Biomedical Books]]|year=2008|isbn=9781604562323|editor-last=Matsumoto|editor-first=Takumi|location=New York|pages=13–30|chapter=Studies on Perilla Relating to Its Essential Oil and Taxonomy|chapter-url=https://books.google.com/books?&id=l5Mo0M5CqQcC&oi=fnd&pg=PA13}}</ref>
The classification of ''Perilla'' is confused, partly because botanists struggled with distinguishing the two distinct [[cultigen]]s as different species or variations.<ref name="Nitta" /><ref name="Yu">{{Cite book|url=https://books.google.com/books?id=M5A71gsWUskC&pg=PA1|title=Perilla: The Genus Perilla|publisher=[[CRC Press]]|year=1997|isbn=9789057021718|editor-last=Yu|editor-first=He-ci|location=London|pages=1–3|editor-last2=Kosuna|editor-first2=Kenichi|editor-last3=Haga|editor-first3=Megumi}}</ref> Until a few decades ago, [[Perilla frutescens var. crispa|''P. frutescens'' var. ''crispa'']] was regarded as a species in its own right, distinct from [[Perilla frutescens|''P. frutescens'']], although it was well established that these types readily cross-pollinate.<ref name="Heibonsha">{{Cite book|title=Sekai dai-hyakka jiten|title-link=Heibonsha World Encyclopedia|publisher=Heibonsha|year=1969|editor-last=Satake|editor-first=Yoshisuke|volume=10|location=Tōkyō|pages=246–207|script-title=ja:世界大百科事典|trans-title=Heibonsha World Encyclopedia|chapter=shiso|script-chapter=ja:しそ|orig-year=1968|editor-last2=Nishi|editor-first2=Sadao|editor-last3=Motoyama|editor-first3=Tekishū}}</ref> An early example of dividing the two cultigens into different species is found in [[Jinzō Matsumura|Matsumura]]'s 1884 nomenclature book, where the synonym ''P. arguta'' [[Benth.|<small>Benth.</small>]] is applied to [[Perilla frutescens var. crispa|''P. frutescens'' var. ''crispa'']], and the synonym ''P. ocymoides'' [[Carl Linnaeus|<small>L.</small>]] was applied to [[Perilla frutescens|''P.'' frutescens]].<ref name="Matsumura">{{Cite book|url=https://archive.org/details/b30093077|title=Nippon shokubutsumeii|last=Matsumura|first=Jinzō|publisher=Z.P. Maruya|year=1884|location=Tōkyō|pages=[https://archive.org/details/b30093077/page/136 136]|script-title=ja:日本植物名彙|trans-title=Nomenclature of Japanese plants in Latin, Japanese and Chinese|author-link=Jinzō Matsumura}}</ref><ref name="Roecklein">{{Cite book|url=https://books.google.com/books?id=C8eRalwPua0C&pg=PA349|title=A Profile of Economic Plants|publisher=[[Transaction Publishers]]|year=1987|isbn=9780887381676|editor-last=Roecklein|editor-first=John C.|location=New Brunswick, U.S.A|pages=349|language=en|editor-last2=Leung|editor-first2=PingSun}}</ref><ref name="Kays">{{Cite book|url=https://books.google.com/books?id=9c7LggdxjBcC&pg=PA677|title=Cultivated Vegetables of the World:: A Multilingual Onomasticon|last=Kays|first=S. J.|publisher=[[Wageningen Academic Publishers]]|year=2011|isbn=9789086861644|location=Wageningen|pages=180–181, 677–678}}</ref><ref name="Blaschek">{{Cite book|url=https://books.google.com/books?id=XGXnVRiu3zoC&pg=PA328|title=Hagers Handbuch der Pharmazeutischen Praxis|publisher=[[Gabler Wissenschaftsverlage]]|year=1998|isbn=9783540616191|editor-last=Blaschek|editor-first=Wolfgang|edition=3|location=Berlin|pages=328–|language=de|editor-last2=Hänsel|editor-first2=Rudolf|editor-last3=Keller|editor-first3=Konstantin|editor-last4=Reichling|editor-first4=Jürgen|editor-last5=Rimpler|editor-first5=Horst|editor-last6=Schneider|editor-first6=Georg}}</ref> The species name ''P. ocymoides'' or ''P. ocimoides'' has historically been used to denote [[Perilla frutescens var. crispa|''P. frutescens'' var. ''crispa'']], especially by the Japanese,{{efn|e.g. occurs in Heibonsha 1964 Encyclopedia, though the genus name is misspelt}}<ref name="Heibonsha" /> therefore it should not be considered an interchangeable synonym for either cultigen. Recent genetic research confirms that the cultigens are of a common gene pool, corroborating the [[taxonomist]]s' claim for consolidating the two crops into one species.<ref name="Ito">{{Cite book|title=Phytochemistry Research Progress|last=Ito|first=Michiho|publisher=[[Nova Biomedical Books]]|year=2008|isbn=9781604562323|editor-last=Matsumoto|editor-first=Takumi|location=New York|pages=13–30|chapter=Studies on Perilla Relating to Its Essential Oil and Taxonomy|chapter-url=https://books.google.com/books?id=l5Mo0M5CqQcC&pg=PA13}}</ref>


Existing ''Perilla species'':
Existing ''Perilla species'':
* [[Perilla frutescens|''P. frutescens'']] – also called Korean perilla or ''deulkkae''; the leaves are called ''kkaennip''
* [[Perilla frutescens|''P. frutescens'']] – also called Korean perilla or ''deulkkae''; the leaves are called ''kkaennip''
* [[Perilla frutescens var. crispa|''P. frutescens'' var. ''crispa'']] – also called ''shiso'', ''tía tô'', or Japanese Basil<ref>{{cite news |last=Mendelson |first=Charlotte |date=29 Jul 2017 |title=The Garden’s Tiny Culinary Transformations |url= https://www.newyorker.com/culture/onward-and-upward-in-the-garden/the-gardens-tiny-culinary-transformations|work=[[The New Yorker]] |location= |access-date=18 Nov 2017 }}</ref>
** [[Perilla frutescens var. crispa|''P. f.'' var. ''crispa'']] – also called ''shiso'', ''tía tô'', or Japanese basil<ref>{{cite magazine |last=Mendelson |first=Charlotte |date=29 Jul 2017 |title=The Garden's Tiny Culinary Transformations |url= https://www.newyorker.com/culture/onward-and-upward-in-the-garden/the-gardens-tiny-culinary-transformations|magazine=[[The New Yorker]] |access-date=18 Nov 2017 }}</ref>
* ''[[Lemon perilla|P. frutescens var. hirtella]]'' – also called lemon perilla<ref name="Seidemann">{{Cite book|url=https://books.google.com/books?id=jjkAHi2IeOkC&pg=PA278|title=World Spice Plants: Economic Usage, Botany, Taxonomy|last=Seidemann|first=Johannes|publisher=[[Springer Science+Business Media|Springer]]|year=2005|isbn=9783540222798|location=Berlin|pages=278}}</ref>
** [[Lemon perilla|''P. f.'' var. ''hirtella'']] – also called lemon perilla<ref name="Seidemann">{{Cite book|url=https://books.google.com/books?id=jjkAHi2IeOkC&pg=PA278|title=World Spice Plants: Economic Usage, Botany, Taxonomy|last=Seidemann|first=Johannes|publisher=[[Springer Science+Business Media|Springer]]|year=2005|isbn=9783540222798|location=Berlin|pages=278}}</ref>
Possible ''Perilla'' species with insufficient description and without known herbarium specimens include:
Possible ''Perilla'' species with insufficient description and without known herbarium specimens include:
* ''P. cavaleriei'' <small>H.Lév.</small>
* ''P. cavaleriei'' <small>H.Lév.</small>
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| width2 = 1932
| width2 = 1932
| height2 = 1239
| height2 = 1239
| caption2 = ''[[Aojiso]]'', the edible leaves of ''[[Perilla frutescens var. crispa]]'' (''P. frutescens'' var. ''crispa'')
| caption2 = ''Aojiso'', the edible leaves of ''[[Perilla frutescens var. crispa]]'' (''P. frutescens'' var. ''crispa'')
}}
}}
* ''P. elata'' {{Au|[[David Don|D.Don.]]}} = ''[[Elsholtzia blanda]]'' {{Au|([[Benth.]]) Benth.}}<ref name="Bentham">{{Cite book|url=https://books.google.com/books?id=uGhTAAAAYAAJ&pg=PA166|title=Labiatarum Genera et Species|last=Bentham|first=George|publisher=James Ridgway and Sons|year=1836|location=London|pages=166|author-link=George Bentham}}</ref>
* ''P. elata'' {{Au|[[David Don|D.Don.]]}} = ''[[Elsholtzia blanda]]'' {{Au|([[Benth.]]) Benth.}}<ref name="Bentham">{{Cite book|url=https://books.google.com/books?id=uGhTAAAAYAAJ&pg=PA166|title=Labiatarum Genera et Species|last=Bentham|first=George|publisher=James Ridgway and Sons|year=1836|location=London|pages=166|author-link=George Bentham}}</ref>
* ''P. fruticosa'' {{Au|D.Don}} = ''[[Elsholtzia fruticosa]]'' {{Au|(D. Don) Rehder}}<ref name="Bentham" /><ref name="Wiersema">{{Cite book|url=http://www.ars-grin.gov/~sbmljw/introd.htm|title=World Economic Plants: A Standard Reference|last=Wiersema|first=John H.|last2=León|first2=Blanca|publisher=[[CRC Press]]|year=1999|isbn=9780849321191}}</ref>
* ''P. fruticosa'' {{Au|D.Don}} = ''[[Elsholtzia fruticosa]]'' {{Au|(D. Don) Rehder}}<ref name="Bentham" /><ref name="Wiersema">{{Cite book|url=http://www.ars-grin.gov/~sbmljw/introd.htm|title=World Economic Plants: A Standard Reference|last1=Wiersema|first1=John H.|last2=León|first2=Blanca|publisher=[[CRC Press]]|year=1999|isbn=9780849321191}}</ref>
* ''P. lanceolata'' {{Au|Benth}} = ''[[Mosla scabra]]'' {{Au|(Thunb.) C.Y.Wu & H.W.Li}}
* ''P. lanceolata'' {{Au|Benth}} = ''[[Mosla scabra]]'' {{Au|(Thunb.) C.Y.Wu & H.W.Li}}
* ''P. leptostachya'' {{Au|D.Don.}} = ''[[Elsholtzia stachyodes]]'' {{Au|(Link) Raizada & H.O.Saxena}}<ref name="Bentham" /><ref name="Pangtey">{{Cite book|title=High altitudes of the Himalaya: Biodiversity, ecology & environment|last=Pangtey|first=Y. P. S.|last2=Rawal|first2=R. S.|publisher=Gyanodaya Prakashan|year=1994|location=Nainital, India|pages=338}}</ref>''
* ''P. leptostachya'' {{Au|D.Don.}} = ''[[Elsholtzia stachyodes]]'' {{Au|(Link) Raizada & H.O.Saxena}}<ref name="Bentham" /><ref name="Pangtey">{{Cite book|title=High altitudes of the Himalaya: Biodiversity, ecology & environment|last1=Pangtey|first1=Y. P. S.|last2=Rawal|first2=R. S.|publisher=Gyanodaya Prakashan|year=1994|location=Nainital, India|pages=338}}</ref>''
* ''P. marathrosma'' {{Au|Spreng.}} = ''[[Agastache foeniculum]]'' {{Au|(Pursh) Kuntze}}
* ''P. marathrosma'' {{Au|Spreng.}} = ''[[Agastache foeniculum]]'' {{Au|(Pursh) Kuntze}}
* ''P. nankinensis'' {{Au|Wender.}} = ''[[Plectranthus scutellarioides]]'' {{Au|(L.) R.Br.}}
* ''P. nankinensis'' {{Au|Wender.}} = ''[[Plectranthus scutellarioides]]'' {{Au|(L.) R.Br.}}
Line 51: Line 46:


==Use==
==Use==
⚫
===Culinary use===

⚫
=== Culinary use ===
{{further|Deulkkae|Shiso|Perilla oil}}
{{further|Deulkkae|Shiso|Perilla oil}}
Perilla varieties are cultivated and consumed mainly in [[Korea]], [[Japan]], [[Thailand]] and [[Vietnam]].<ref>{{Cite conference|url=http://www.tropentag.de/2016/proceedings/proceedings.pdf|last1=Grbic|first1=Nikolina|last2=Pinker|first2=Ina|last3=Paschko|first3=Kerstin|last4=Böhme|first4=Michael|date=18–21 September 2016|title=The Nutritional Treasure of Leafy Vegetables-Perilla frutescens|editor-last1=Freyer|editor-first1=Bernhard|editor-last2=Tielkes|editor-first2=Eric|book-title=Solidarity in a competing world - fair use of resources|conference=Tropentag 2016 Wien: International Research on Food Security, Natural Resource Management and Rural Development|conference-url=http://www.tropentag.de|publication-place=Göttingen, Germany|publisher=Cuvillier|page=105|isbn=978-3-9801686-7-0|access-date=5 May 2017}}</ref> ''[[Perilla frutescens|P. frutescens]]'' leaves, seeds, and [[perilla oil|seed oil]] are used extensively in [[Korean cuisine]], while [[Perilla frutescens var. crispa|''P. frutescens'' var. ''crispa'']] leaves, seeds, and sprouts are used in [[Japanese cuisine|Japanese]] and [[Vietnamese cuisine]]s.<ref>{{Cite book|title=New crops|last=Brenner|first=David M.|publisher=[[John Wiley & Sons]]|year=1993|isbn=0-471-59374-5|editor-last=J.|editor-first=Janick|location=New York|pages=322–328|chapter=Perilla: Botany, Uses and Genetic Resources|editor-last2=Simon|editor-first2=J. E.|chapter-url=https://hort.purdue.edu/newcrop/proceedings1993/V2-322.html}}</ref>
Perilla varieties are cultivated and consumed mainly in [[Korea]], [[Japan]], [[Thailand]] and [[Vietnam]].<ref>{{Cite conference|url=http://www.tropentag.de/2016/proceedings/proceedings.pdf|last1=Grbic|first1=Nikolina|last2=Pinker|first2=Ina|last3=Paschko|first3=Kerstin|last4=Böhme|first4=Michael|date=18–21 September 2016|title=The Nutritional Treasure of Leafy Vegetables-Perilla frutescens|editor1-last=Freyer|editor1-first=Bernhard|editor2-last=Tielkes|editor2-first=Eric|book-title=Solidarity in a competing world - fair use of resources|conference=Tropentag 2016 Wien: International Research on Food Security, Natural Resource Management and Rural Development|conference-url=http://www.tropentag.de|location=Göttingen, Germany|publisher=Cuvillier|page=105|isbn=978-3-9801686-7-0|access-date=5 May 2017}}</ref> ''[[Perilla frutescens|P. frutescens]]'' leaves, seeds, and [[perilla oil|seed oil]] are used extensively in [[Korean cuisine]], while [[Perilla frutescens var. crispa|''P. frutescens'' var. ''crispa'']] leaves, seeds, and sprouts are used in [[Japanese cuisine|Japanese]] and [[Vietnamese cuisine]]s.<ref>{{Cite book|title=New crops|last=Brenner|first=David M.|publisher=[[John Wiley & Sons]]|year=1993|isbn=978-0-471-59374-4|editor-last=J.|editor-first=Janick|location=New York|pages=[https://archive.org/details/newcrops0000nati/page/322 322–328]|chapter=Perilla: Botany, Uses and Genetic Resources|editor-last2=Simon|editor-first2=J. E.|chapter-url=https://hort.purdue.edu/newcrop/proceedings1993/V2-322.html|url=https://archive.org/details/newcrops0000nati/page/322}}</ref>


== Notes ==
===Herbalism===
Perilla is one of the 50 fundamental herbs in [[Traditional Chinese Medicine]]. It is called "zi su" (紫苏/紫蘇), and is used to disperse wind-cold, bloating, and stomach and lung problems. It is sometimes paired with [[huo xiang| Tu Huo Xiang]] or [[Patchouli| Guang Huo Xiang]] to dispel dampness and tonify [[qi]].

==Notes==
{{notelist}}
{{notelist}}


==References==
==References==
{{Reflist|30em}}
{{Reflist|30em}}
⚫
{{Commons category|Perilla}}


{{Authority control}}
{{Authority control}}
⚫
{{Commons category|Perilla}}

{{Taxonbar|from=Q7168518}}
{{Taxonbar|from=Q7168518}}


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[[Category:Lamiaceae genera]]
[[Category:Lamiaceae genera]]
[[Category:Taxa named by Carl Linnaeus]]
[[Category:Taxa named by Carl Linnaeus]]
[[Category:Lamiaceae]]

Latest revision as of 19:14, 31 August 2023

For other uses, see Perilla (disambiguation).

Perilla
Perilla frutescens
Scientific classification Edit this classification
Kingdom: Plantae
Clade: Tracheophytes
Clade: Angiosperms
Clade: Eudicots
Clade: Asterids
Order: Lamiales
Family: Lamiaceae
Subfamily: Nepetoideae
Tribe: Elsholtzieae
Genus: Perilla
L.

Perilla is a genus consisting of one major Asiatic crop species Perilla frutescens and a few wild species in nature belonging to the mint family, Lamiaceae. The genus encompasses several distinct varieties of Asian herb, seed, and vegetable crop, including P. frutescens (deulkkae) and P. frutescens var. crispa (shiso).[1] The genus name Perilla is also a frequently employed common name ("perilla"), applicable to all varieties.[2][3] Perilla varieties are cross-fertile and intra-specific hybridization occurs naturally.[1] Some varieties are considered invasive.[4]

Taxa and synonyms[edit]

The classification of Perilla is confused, partly because botanists struggled with distinguishing the two distinct cultigens as different species or variations.[1][5] Until a few decades ago, P. frutescens var. crispa was regarded as a species in its own right, distinct from P. frutescens, although it was well established that these types readily cross-pollinate.[6] An early example of dividing the two cultigens into different species is found in Matsumura's 1884 nomenclature book, where the synonym P. arguta Benth. is applied to P. frutescens var. crispa, and the synonym P. ocymoides L. was applied to P. frutescens.[7][8][9][10] The species name P. ocymoides or P. ocimoides has historically been used to denote P. frutescens var. crispa, especially by the Japanese,[a][6] therefore it should not be considered an interchangeable synonym for either cultigen. Recent genetic research confirms that the cultigens are of a common gene pool, corroborating the taxonomists' claim for consolidating the two crops into one species.[11]

Existing Perilla species:

Possible Perilla species with insufficient description and without known herbarium specimens include:

  • P. cavaleriei H.Lév.
  • P. heteromorpha Carrière
  • P. setoyensis G.Honda

The past legacies and subsequent reclassification of taxa has led to confusion of nomenclature.[14][5] The red or purple leafed variety of P. frutescens var. crispa had been dubbed P. nankinensis,[5] and this label was used throughout the 19th century in the West following the introduction of the species for ornamental planting. Whether green-leafed or red-leafed, the perillaldehyde factor that characterizes the unique P. frutescens var. crispa fragrance may turn out to be present or absent in the individual or population, and this is not differentiable from outward appearance alone. Chemical studies classify the genus into different chemotypes, depending on the essential oils they contain. Three wild species that are endemic to Japan are recognized as genetically distinct from the cultivated P. frutescens var. crispa,[11] however, some references treat, e.g., P. frutescens var. hirtella as the same species as the P. frutescens var. crispa.[10]

Formerly placed here[edit]

Kkaennip, the edible leaves of Perilla frutescens (P. frutescens)
Aojiso, the edible leaves of Perilla frutescens var. crispa (P. frutescens var. crispa)

Use[edit]

Culinary use[edit]

Further information: Deulkkae, Shiso, and Perilla oil

Perilla varieties are cultivated and consumed mainly in Korea, Japan, Thailand and Vietnam.[18] P. frutescens leaves, seeds, and seed oil are used extensively in Korean cuisine, while P. frutescens var. crispa leaves, seeds, and sprouts are used in Japanese and Vietnamese cuisines.[19]

Herbalism[edit]

Perilla is one of the 50 fundamental herbs in Traditional Chinese Medicine. It is called "zi su" (紫苏/紫蘇), and is used to disperse wind-cold, bloating, and stomach and lung problems. It is sometimes paired with Tu Huo Xiang or Guang Huo Xiang to dispel dampness and tonify qi.

Notes[edit]

  1. ^ e.g. occurs in Heibonsha 1964 Encyclopedia, though the genus name is misspelt

References[edit]

  1. ^ a b c Nitta, Miyuki; Lee, Ju Kyong; Ohnishi, Ohmi (2003). "AsianPerilla crops and their weedy forms: Their cultivation, utilization and genetic relationships". Economic Botany. 57 (2): 245–253. doi:10.1663/0013-0001(2003)057[0245:APCATW]2.0.CO;2. ISSN 0013-0001. S2CID 30209741.
  2. ^ Vaughan, John; Geissler, Catherine (2009). The New Oxford Book of Food Plants (2nd ed.). Oxford: Oxford University Press. p. 340. ISBN 9780199549467.
  3. ^ Staples, George W.; Kristiansen, Michael S. (1999). Ethnic Culinary Herbs: A Guide to Identification and Cultivation in Hawaiʻi. Honolulu: University of Hawaiʻi Press. p. 82. ISBN 9780824820947.
  4. ^ "Beefsteak Plant (Perilla frustescens)". National Park Service. U. S. Department of the Interior. 11 November 2010. Retrieved 4 December 2016.
  5. ^ a b c Yu, He-ci; Kosuna, Kenichi; Haga, Megumi, eds. (1997). Perilla: The Genus Perilla. London: CRC Press. pp. 1–3. ISBN 9789057021718.
  6. ^ a b Satake, Yoshisuke; Nishi, Sadao; Motoyama, Tekishū, eds. (1969) [1968]. "shiso" しそ. Sekai dai-hyakka jiten 世界大百科事典 [Heibonsha World Encyclopedia]. Vol. 10. Tōkyō: Heibonsha. pp. 246–207.
  7. ^ Matsumura, Jinzō (1884). Nippon shokubutsumeii 日本植物名彙 [Nomenclature of Japanese plants in Latin, Japanese and Chinese]. Tōkyō: Z.P. Maruya. pp. 136.
  8. ^ Roecklein, John C.; Leung, PingSun, eds. (1987). A Profile of Economic Plants. New Brunswick, U.S.A: Transaction Publishers. p. 349. ISBN 9780887381676.
  9. ^ Kays, S. J. (2011). Cultivated Vegetables of the World:: A Multilingual Onomasticon. Wageningen: Wageningen Academic Publishers. pp. 180–181, 677–678. ISBN 9789086861644.
  10. ^ a b Blaschek, Wolfgang; Hänsel, Rudolf; Keller, Konstantin; Reichling, Jürgen; Rimpler, Horst; Schneider, Georg, eds. (1998). Hagers Handbuch der Pharmazeutischen Praxis (in German) (3 ed.). Berlin: Gabler Wissenschaftsverlage. pp. 328–. ISBN 9783540616191.
  11. ^ a b Ito, Michiho (2008). "Studies on Perilla Relating to Its Essential Oil and Taxonomy". In Matsumoto, Takumi (ed.). Phytochemistry Research Progress. New York: Nova Biomedical Books. pp. 13–30. ISBN 9781604562323.
  12. ^ Mendelson, Charlotte (29 Jul 2017). "The Garden's Tiny Culinary Transformations". The New Yorker. Retrieved 18 Nov 2017.
  13. ^ Seidemann, Johannes (2005). World Spice Plants: Economic Usage, Botany, Taxonomy. Berlin: Springer. p. 278. ISBN 9783540222798.
  14. ^ Zeevaart, Jan A. D. (1969). "Perilla". In Evans, L. T. (ed.). The Induction of Flowering: Some Case Histories. Ithaca, NY: Cornell University Press. pp. 116–155. ISBN 9780333101537.
  15. ^ a b c d Bentham, George (1836). Labiatarum Genera et Species. London: James Ridgway and Sons. p. 166.
  16. ^ Wiersema, John H.; León, Blanca (1999). World Economic Plants: A Standard Reference. CRC Press. ISBN 9780849321191.
  17. ^ Pangtey, Y. P. S.; Rawal, R. S. (1994). High altitudes of the Himalaya: Biodiversity, ecology & environment. Nainital, India: Gyanodaya Prakashan. p. 338.
  18. ^ Grbic, Nikolina; Pinker, Ina; Paschko, Kerstin; Böhme, Michael (18–21 September 2016). "The Nutritional Treasure of Leafy Vegetables-Perilla frutescens" (PDF). In Freyer, Bernhard; Tielkes, Eric (eds.). Solidarity in a competing world - fair use of resources. Tropentag 2016 Wien: International Research on Food Security, Natural Resource Management and Rural Development. Göttingen, Germany: Cuvillier. p. 105. ISBN 978-3-9801686-7-0. Retrieved 5 May 2017.
  19. ^ Brenner, David M. (1993). "Perilla: Botany, Uses and Genetic Resources". In J., Janick; Simon, J. E. (eds.). New crops. New York: John Wiley & Sons. pp. 322–328. ISBN 978-0-471-59374-4.
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